71peer-reviewed papers
3,664citations
33H-index
54i10-index
14+co-author countries

71 peer-reviewed scientific papers, 2 government reports, 1 book chapter, 3 manuscripts in review and 4 in prep. Collaborators from >14 countries including Australia, USA, Canada, United Kingdom, Denmark, Italy, Brazil, UAE, New Zealand, Saudi Arabia.

Asterisk (*) highlights undergraduate and graduate students at time of project completion.

View on Google Scholar (most up-to-date)

Book Chapters

  1. Esbaugh AJ, *Khursigara AJ, Johansen JL (2018) Toxicity in Aquatic Environments: The Cocktail Effect. In “Development and Environment” (editor Burggren W), Springer Publishers. p 203-234.

International Refereed Papers

  1. *Souza T, Capistran MME, Crowder L, Johansen JL (2026) Herbivores in hot water: thermal limits and avoidance of marine heatwave conditions by Acanthurus triostegus. Coral Reefs, 45(3), 1393-1405. DOI
  2. Steffensen JF, Johansen JL, Domenici P (2026) An integrative view of fish swimming kinematics and energetics: a tribute to 20+ years of teaching and research at Friday Harbor Laboratories. Journal of Experimental Biology, 229(7), jeb251814
  3. Brijs J, *Tran LL, *Moore C, *Souza T, *Schakmann M, *Grellman K, Johansen JL (2025) Outlasting the heat: collapse of herbivorous fish control of invasive algae during marine heatwaves. Global Change Biology, 31(8), p.e70438. DOI
  4. Domenici P, Johansen JL, Karoliussen R, Leggieri F, Dykstra P, Jourdain E (2025) Spatially coordinated predation with division of labor increases feeding success in killer whales. Current Biology, 35(14), 3507-3514.e5
  5. Vaughan GO, Ripley DM, Mitchell MD, McParland D, Johansen JL, Shiels HA, Burt JA (2025) Narrow Margins: Aerobic Performance and Temperature Tolerance of Coral Reef Fishes Facing Extreme Thermal Variability. Global Change Biology, 31(3), p.e70100
  6. Brijs J, *Moore C, *Schakmann M, *Souza T, *Grellman K, *Tran LL, *Patton PT, Johansen JL (2025) Eat more, often: The capacity of piscivores to meet increased energy demands in warming oceans. Science of The Total Environment, 973, p.179105
  7. *Souza T, Brijs J, *Tran L, Crowder L, Johansen JL (2025) Herbivore functions in the hot-seat: Resilience of Acanthurus triostegus to marine heatwaves. PloS one, 20(1), p.e0318410
  8. *Tiddy IC, *Neill CM, *Rosén A, *Hasegawa Y, Domenici P, Johansen JL, Steffensen JF (2025) Effects of social environment and energy efficiency on preferred swim speed in a marine generalist fish, pile perch (Phanerodon vacca). Journal of Experimental Biology, 228(5)
  9. Johansen JL, Mitchell MD, *Vaughan GO, *Ripley DM, Shiels HA, Burt JA (2024) Impacts of ocean warming on fish size reductions on the world’s hottest coral reefs. Nature Communications 15(1), p.5457. DOI
  10. *Frank LC, *Prescott LA, *Scott ME, Domenici P, Johansen JL, Steffensen JF (2024) The effect of progressive hypoxia on swimming mode and oxygen consumption in the pile perch, Phanerodon vacca. Frontiers in Fish Science, 2, 1289848
  11. *Tran LL, Johansen JL (2023) Seasonal variability in resilience of a coral reef fish to marine heatwaves and hypoxia. Global Change Biology, 29(9), 2522-2535
  12. *Kimura H, *Pfalzgraff T, *Levet M, *Kawabata Y, Steffensen JF, Johansen JL, Domenici P (2022) Escaping from multiple visual threats: modulation of escape responses in Pacific staghorn sculpin (Leptocottus armatus). J Exp Biol 225(9)
  13. Nadler LE, McCormick MI, Johansen JL, Domenici P (2021) Social familiarity improves fast-start escape performance in schooling fish. Communications Biology 4(1), 1-10
  14. Brijs J, Føre M, Gräns A, Clark TD, Axelsson M, Johansen JL (2021) Bio-sensing technologies in aquaculture: how remote monitoring can bring us closer to our farm animals. Philosophical Transactions of the Royal Society B 376(1830), 20200218
  15. *Khursigara AJ, *Rowsey LE, Johansen JL, Esbaugh AJ (2021) Behavioral changes in a coastal marine fish lead to increased predation risk following oil exposure. Environ. Sci. Technol. 55(12), 8119-8127
  16. *Nay T, Hoey AS, Rummer JL, Steffensen JF, Johansen JL (2021) Species interactions alter the selection of thermal environment in a coral reef fish. Oecologia, 1-9
  17. *Nay TJ, Longbottom RJ, *Gervais CR, Johansen JL, Steffensen JF, Rummer JL, Hoey AS (2021) Regulate or tolerate: Thermal strategy of a coral reef flat resident, the epaulette shark, Hemiscyllium ocellatum. Journal of Fish Biology 98(3), 723-732
  18. *Schakmann M, *Becker V, *Soegaard M, Johansen JL, Steffensen JF, Domenici P (2021) Latency of mechanically stimulated escape responses in the Pacific spiny dogfish, Squalus suckleyi. J Exp Biol 224(3), jeb230698
  19. Johansen JL, Nadler LE, *Habary A, *Bowden AJ, Rummer JL (2021) Thermal acclimation of tropical fishes to marine heat waves. eLife 10, e59162
  20. *Khursigara AJ, Johansen JL, Esbaugh AJ (2021) The effects of acute crude oil exposure on growth and competition in the red drum, Sciaenops ocellatus. Science of the Total Environment 751, 141804
  21. Johansen JL, Akanyeti O, Liao JC (2020) Oxygen consumption of drift-feeding rainbow trout: the energetic tradeoff between locomotion and feeding in flow. J Exp Biol
  22. Johansen JL†, Brandl SJ†, Casey JM, Tornabene L, Morais RA, Burt JA (2020) Extreme environmental conditions reduce coral reef fish biodiversity and productivity. Nature Communications 11(1), 1-14 († shared first authorship)
  23. Burt JA, Camp EF, Enochs IC, Johansen JL, Morgan MS, Riegl B, Hoey AS (2020) Insights from extreme coral reefs in a changing world. Coral Reefs
  24. *Nay TJ, Johansen JL, Rummer JL, Steffensen JF, Pratchett MS, Hoey AS (2020) Habitat complexity influences selection of thermal environment in a common coral reef fish. Conservation Physiology 8(1), coaa070
  25. *Luongo SM, *Ruth A, *Gervais CR, Korsmeyer KE, Johansen JL, Domenici P, Steffensen JF (2020) Bidirectional cyclical flows increase energetic costs of station holding for a labriform swimming fish, Cymatogaster aggregata. Conservation Physiology 8(1), coaa077
  26. McHenry MJ, Johansen JL, *Soto AP, *Free BA, *Paley DA, Liao JC (2019) The pursuit strategy of predatory bluefish (Pomatomus saltatrix). Proceedings of the Royal Society B 286
  27. *Rowsey LE, Johansen JL, *Khursigara AJ, Esbaugh AJ (2019) Oil exposure impairs predator-prey dynamics in larval red drum (Sciaenops ocellatus). Marine and Freshwater Research Special Edition (70th anniversary)
  28. Svendsen MBS, Johansen JL, Bushnell PG, Skov PV, Norin T, Domenici P, Steffensen JF, Abe A (2019) Are all bony fishes oxygen regulators? Evidence for oxygen regulation in a putative oxygen conformer, the swamp eel Synbranchus marmoratus. Journal of Fish Biology 94, 178-182
  29. Johansen JL, Esbaugh AJ (2019) Oil-induced responses of cardiac and red muscle mitochondria in red drum (Sciaenops ocellatus). Comparative Biochemistry and Physiology Part C, 219, 35-41
  30. *Khursigara AJ, Johansen JL, Esbaugh AJ (2018) Social competition in red drum (Sciaenops ocellatus) is influenced by crude oil exposure. Aquatic Toxicology 203, 194-201
  31. *Nay TJ, *Gervais CR, Hoey AS, Johansen JL, Steffensen JF, Rummer JL (2018) The emergence emergency: A mudskipper’s response to temperatures. Journal of Thermal Biology 78, 65-72
  32. *Gervais CR, *Nay TJ, effects of elevated temperatures in a benthic elasmobranch, Hemiscyllium ocellatum. Marine Biology 165(11)
  33. *Christensen EAF, *Illing B, *Iversen NS, Johansen JL, Domenici P, Steffensen JF (2018) Effects of salinity on swimming performance and oxygen consumption rate of shiner perch Cymatogaster aggregata. J Exp Biol 504, 32-37
  34. *Pan YK, *Khursigara AJ, Johansen JL, Esbaugh AJ (2018) The effects of oil induced respiratory impairment on two indices of hypoxia tolerance in Atlantic croaker (Micropogonias undulatus). Chemosphere 200, 143-150
  35. *van der Hoop JM, *Byron M, *Ozolina K, Johansen JL, Domenici P, Steffensen JF (2018) Turbulent flow reduces oxygen consumption in the labriform swimming shiner perch, Cymatogaster aggregata. J Exp Biol 221(11)
  36. He S, Johansen JL, Hoey AS, Pappas MK, Berumen ML (2017) Molecular confirmation of hybridization between Dascyllus reticulatus × Dascyllus aruanus from the Great Barrier Reef. Marine Biodiversity 49, 395-404
  37. Johansen JL, Allan BJM, Rummer JL, Esbaugh AJ (2017) Oil exposure disrupts early life-history stages of coral reef fishes. Nature Ecology and Evolution 1, 1146-1152
  38. Ern R, Johansen JL, Rummer JL, Esbaugh AJ (2017) Effects of hypoxia and ocean acidification on the upper thermal niche boundaries of coral reef fishes. Biology Letters 13(7), 20170135
  39. Stewart WJ, Johansen JL, Liao JC (2017) A non-toxic dose of Cobalt Chloride blocks hair cells of the zebrafish lateral line. Hearing Research 350, 17-21
  40. Johansen JL, Esbaugh AJ (2017) Sustained impairment of respiratory function and swim performance following acute oil exposure in a coastal marine fish. Aquatic Toxicology 187, 82-89
  41. *Shi X, *Moller JS, *Hojgaard J, Johansen JL, Steffensen JF, Liu D, Domenici P (2017) The angular position of a refuge affects escape responses in staghorn sculpin (Leptocottus armatus). Journal of Fish Biology 90(6), 2434-2442
  42. Johansen JL, He S, Frank G, Pappas MK, Berumen ML, Hoey AS (2017) Hybridization between the benthic nesting and pair spawning damselfishes Dascyllus aruanus and D. reticulatus. Coral Reefs 1-1
  43. *Mazloumi N, Johansen JL, Doubleday Z, Gillanders BW (2017) Q10 measures of metabolic performance and critical swimming speed in a temperate fish (King George whiting, Sillaginodes punctatus). Journal of Fish Biology 90(5), 2200-2205
  44. *Habary A, Johansen JL, *Nay TJ, Steffensen JF, Rummer JL (2017) Adapt, move, or die — how will tropical coral reef fishes cope with ocean warming? Global Change Biology 23(2), 566-577
  45. Hoey AS, Howells E, Johansen JL, Hobbs JPA, Messmer V, McCowan DM, Wilson SK, Pratchett MS (2016) Recent advances in understanding the effects of climate change on coral reefs. Diversity 8(12)
  46. Brooker RM, Feeney WE, White JR, Manassa RP, Johansen JL, Dixson DL (2016) Using insights from animal behaviour and behavioural ecology to inform marine conservation initiatives. Animal Behavior 120, 211-221
  47. Rummer JL, Binning SA, Roche DG, Johansen JL (2016) Methods matter: Considering technique when estimating metabolic rates of fishes. Conservation Physiology 4
  48. *Snyder S, *Nadler LE, *Bayley JS, *Svendsen MBS, Johansen JL, Domenici P, Steffensen JF (2016) Effect of closed v. intermittent-flow respirometry on hypoxia tolerance in the shiner perch Cymatogaster aggregata. Journal of Fish Biology 88, 252-264
  49. Johansen JL, Pratchett MS, Messmer V, Coker DJ, Tobin AJ, Hoey AS (2015) Large predatory coral trout species unlikely to meet increasing energetic demands in a warming ocean. Scientific Reports 5, 13830
  50. Johansen JL, Steffensen JF, Jones GP (2015) Winter temperatures decrease swimming performance and limit distributions of tropical damselfishes. Conservation Physiology 3, cov039
  51. Ferrari MCO, McCormick MI, Allen BJM, Choi R, Ramasamy RA, Johansen JL, Mitchell MD, Chivers DP (2015) Living in a risky world: the onset and ontogeny of an integrated antipredator phenotype in a coral reef fish. Scientific Reports 5, 15537
  52. *Nay TJ, Johansen JL, *Habary A, Steffensen JF, Rummer JL (2015) Behavioural thermoregulation in a temperature-sensitive coral reef fish. Coral Reefs
  53. Domenici P, Norin T, Bushnell PG, Johansen JL, Skov PV, Steffensen JF (2015) Fast-starting for a breath: Air breathing in Hoplosternum littorale. Biology Open
  54. Johansen JL, Messmer V, Coker DJ, Hoey A, Pratchett M (2014) Increasing ocean temperatures reduce activity patterns of a large commercially important coral reef fish. Global Change Biology 20, 1067-1074
  55. Johansen JL (2014) Quantifying water flow within aquatic ecosystems using load cell sensors: A profile of currents experienced by coral reef organisms around Lizard Island, Great Barrier Reef, Australia. PloS One, 9(1), e83240
  56. *Roche DG, *Taylor MK, *Binning SA, Johansen JL, Steffensen JF, Domenici P (2014) Unsteady flow affects swimming energetics in a labriform fish (C. aggregata). J Exp Biol 217, 414-422
  57. *Lucon-Xiccato T, *Nati JJH, *Blasco FR, Johansen JL, Steffensen JF, Domenici P (2014) Severe hypoxia impairs lateralization in a marine teleost fish. J Exp Biol 217, 4115-4118
  58. Johansen JL and Jones GP (2013) Sediment induced turbidity impairs foraging performance and prey choice in planktivorous coral reef fishes. Ecological Applications 23(6), 1504
  59. Roche DG, Binning SA, Bosiger Y, Johansen JL, Rummer JL (2013) Finding the best estimates of metabolic rates in a coral reef fish. J Exp Biol
  60. Fulton CJ, Johansen JL, Steffensen JF (2013) Energetic extremes in coral reef fishes. PLoS One 8(1), e54033
  61. *Wenger AS, Johansen JL, McCormick M, Jones GP (2012) Increasing suspended sediment reduces foraging, growth and condition of a planktivorous damselfish. JEMBE 428, 43-48
  62. Johansen JL, Jones GP (2011) Increasing ocean temperature reduces the metabolic performance and swimming ability of coral reef fishes. Global Change Biology 17(9), 2971-2979. → Nature Climate Change hotspot feature
  63. *Wenger AS, Johansen JL, Jones GP (2011) Suspended sediment impairs habitat choice and olfactory discrimination in two coral reef fishes. Coral Reefs 30(4), 879-887
  64. Johansen JL, Vaknin R, Domenici P, Steffensen JF (2010) Energetic and Kinematic benefits of schooling in Labriform fishes. Marine Ecology Progress Series 420, 221-229
  65. Johansen JL, Bellwood DR, Fulton CJ (2008) Coral reef fishes exploit flow refuges in high-flow habitats. Marine Ecology Progress Series 360, 219-226
  66. Johansen JL, Fulton CJ, Bellwood DR (2007) Avoiding the flow: refuges expand the swimming potential of coral reef fishes. Coral Reefs 26(3), 577-583. Virginia Chadwick Award for Excellence in Scientific Publishing
  67. Johansen JL, Fulton CJ, Bellwood DR (2007) Estimating the sustained swimming ability of coral reef fishes. Marine and Freshwater Research 58(3), 233-239. Front cover paper
  68. Johansen JL, Herbert NA, Steffensen JF (2006) The behavioural and physiological response of Atlantic cod Gadus morhua L. to short-term acute hypoxia. Journal of Fish Biology 68, 1918

Government Reports

  1. Hoey AS, Pratchett MS, Johansen JL, Hoey J (2014) Marine ecological survey of Elizabeth and Middleton reefs, Lord Howe Commonwealth Marine Reserve. The Department of the Environment, Australian Government.
  2. Woodcock SH, Johansen JL, Steer MA, Gaylard SG, Prowse TAA, Gillanders BM (2014) Regional Sustainability Planning in the Upper Spencer Gulf — Investigating potential impacts of shipping on Giant Australian cuttlefish. The Department of the Environment, Australian Government.